All Angles Creatures
Squamata · AgamidaeEntry 15,539 of 21,572
Blacktail Toadhead Agama, Phrynocephalus maculatus

Reptilia · Squamata

Phrynocephalus maculatusAnderson, 1872

Blacktail Toadhead Agama

Photograph by Mohammad Amin Ghaffari, CC BY, via iNaturalist

A ground-dwelling, day-active, omnivorous lizard from the Saharo-Sindian region, reaching 91 mm snout to vent.

Also called Spotted Toad-headed Agama.

ReptiliaSquamataAgamidaePhrynocephalus

Grows to
91mm
greatest recorded
Eats
Omnivorous
Active
Diurnal
Lives
Terrestrial
Body length across Agamidae

Longer than only 49% of all agamids anyone has measured.

49% of 583 species are smaller51% larger
32 mmthe whole family, smallest to largest418 mm

The bar is the rank itself: its length is the share of the family this animal exceeds. The chart below is the family’s real spread, on a logarithmic scale because body length here runs from 32 to 418 mm. The solid rule is this animal. Every bar is a count of real species, and nothing on this figure is estimated.

What is known

35 sourced facts, each taken from a named scientific dataset. How to read these.

Cited from a dataset unless a row is marked otherwise.

Standing in the world

Still living
Extant
Conservation status
Least Concern
Status assessed · the year the Red List last looked
2012
Population trend
Stable

The animal itself

Limbs
Four legs
Venomous
No
Fangs
None
Body temperature when active · measured in the wild
38.8°C102 °F

mean body temperature of active individuals in nature

Body temp in the wild · mean, when active
38.8°C

body temperature of active animals in nature

Size

Length snout to vent · greatest recorded
91mm3.5 in

max 91 mm SVL (Feldman et al. 2016)

Length snout to vent · adult female mean
61.8mm2.5 in
Largest female · snout to vent
61.8mm2.5 in

snout-vent length; the source column mixes this with straight carapace length

Body mass · greatestcomputed
24.94g0.88 oz

computed from length by a family-level allometric equation, not weighed

How it lives

Diet
Omnivorous

insects (e.g., Hojati et al. 2014: "No plant consumption was observed in this species.")/ Omnivorous (Greene 1982, Rodda 2020) / arachnids, plants and insects (Kamali 2020)

How it hunts
Sit and wait

e.g., Gardner 2013

Active
Diurnal

e.g., Gardner 2013, Hojati et al. 2014, Kamali 2020, Albarwari 2021

Where it lives
Terrestrial

e.g., Kamali 2020

Habitat
Desert, Grassland
Biome
Deserts & Xeric Shrublands
Active season · months of the year
5.5–12months
Region
Saharo-Sindian
Mainland or island
Mainland
Elevation · above sea level
10–3000m

500-3000 (Hojati et al. 2014), 2-10 (Al-Barazengy 2015), 35 m (Mohammed et al. 2015), 100-3000 (Albarwari 2021)

Range size
2,828,671km²
Home range · largest recorded
3297.5m2

Breeding

Reproduction
Oviparous
Average clutch
2.5eggs
Eggs or young per clutch
1–4eggs

2-8 (Albarwari 2021; doubtful - before all I had was 1-4)

Naming and history

Suborder
Sauria
Infraorder
Acrodontia
Subspecies
Phrynocephalus maculatus longicaudatus HAAS 1957 Phrynocephalus maculatus maculatus ANDERSON 1872
Described by
Anderson
Moved to another genus · since it was first described
No
Described in
1872
Type locality
Iran

Keeping one

14.5 in × 14.5 in × 9 in(36 × 36 × 23 cm)

Minimum length × width × height, for one animal. Bigger is always better.

The standard sizes ground-dwelling species at 4 × 4 × 2.5 times snout-vent length.

Mindestanforderungen an die Haltung von Reptilien (Sachverständigengutachten), German Federal Ministry for Nutrition, Agriculture and Forestry, Animal Welfare Division, 10 January 1997. The standard states these floor areas are recommendations rather than binding limits.

From this animal’s own snout-vent length of 3.5 in (91 mm), at 4 × 4 × 2.5 times it, following the German federal welfare standard — how this is worked out.

Active animals measured in the wild averaged 102 °F (38.8 °C).

Mean body temperature of active individuals measured in nature. A measurement of the animal, not a setting for a thermostat.

Feeding

A prey item should be no wider than the space between the animal's eyes.

Widely used across reptile husbandry references. The stated reason is that the gap approximates the width of the throat, so wider prey is associated with impaction. This is established practice, not a measurement.

3kJ per day

Worked out for an animal of 25 g, the heaviest recorded for Blacktail Toadhead Agama — so it is the top of the range, not a figure for a particular animal. Weigh yours and the number will be lower.

An upper estimate, not a feeding target, and not veterinary advice. The equation covers only what an animal burns staying alive, so a growing, gravid or recovering animal needs more — and it was measured on wild animals, which move far more than one in an enclosure. Body condition and a vet who knows the species are what a feeding decision should rest on. Where this comes from.

What nobody knows

These are not gaps in our research. No scientific dataset records them for this species — and saying so is information you cannot get from a reference that simply leaves a field out.

  • Length snout to vent · adult male mean
  • Length snout to vent · adult mean
  • Length nose to tail · greatest recorded
  • Female length · mean, nose to tail
  • Length nose to tail · adult male mean
  • Length nose to tail · adult mean
  • Largest male · snout to vent
  • Largest female · shell length
  • Largest male · shell length
  • Hatchling length · snout to vent
  • Hatchling length · nose to tail
  • Largest juvenile · snout to vent
  • Largest juvenile · shell length
  • Egg length
  • Egg width
  • Body mass · a typical adult
  • Body mass · adult female, mean
  • Body mass at hatching
  • Longest recorded life
  • Preferred body temperature · chosen in a laboratory gradient
  • Eats arthropods
  • Eats vertebrates
  • Eats plant matter
  • How the young develop
  • Gestation or incubation
  • Mean clutch · smallest reported
  • Mean clutch · largest reported
  • Clutches in a year
  • Clutches a year · fewest
  • Clutches a year · most
  • First breeds from · youngest recorded
  • First breeds by · oldest recorded
  • Hatchling length · smallest recorded
  • Hatchling length · largest recorded
  • Age at first breeding · female
  • Mature from · youngest recorded
  • Mature by · oldest recorded
  • Size at maturity · smallest recorded
  • Size at maturity · largest recorded
  • Colour on the back
  • Pattern on the back
  • Pupil
  • Sex decided by
  • Red List criteria
  • Density · lowest recorded
  • Density · highest recorded
  • Active in

Others in Agamidae

Take this data

Everything on this page is available as data under CC BY 4.0 — use it freely, credit it, no permission needed.

This species as JSON · the whole dataset

Cite this page

The Reptile & Amphibian Encyclopedia (2026). Phrynocephalus maculatus. Retrieved 2026-09-08, from https://www.allanglescreatures.com/species/phrynocephalus-maculatus

@misc{phrynocephalus_maculatus_2026,
  title  = {Phrynocephalus maculatus},
  author = {{The Reptile & Amphibian Encyclopedia}},
  year   = {2026},
  url    = {https://www.allanglescreatures.com/species/phrynocephalus-maculatus}
}

Sources

  1. Meiri, S. (2024). SquamBase — a database of squamate (Reptilia: Squamata) traits. Global Ecology and Biogeography. Public domain (CC0).
  2. Oskyrko, O. et al. (2024). ReptTraits: a comprehensive dataset of ecological traits in reptiles. Scientific Data. Used under CC BY 4.0.
  3. Catalogue of Life (dataset 3LR), used under CC BY.